The FullName field on the Person document contains the Notes username. When an email is sent, Crossware Mail Signature looks up the sender of the email in the Domino address book, obtains the FullName, and uses it when processing signature configurations that are applied to groups. It will check the group's members to see if that user is a member.
For more information on how Crossware Mail Signature processes the sender of emails, see How the sender of an email is determined.
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CMS DOM v4